Cephennodes er en slekt av små biller som nå blir regnet til familien kortvinger (Staphylinidae). De er rovdyr som har spesialisert seg på midd. Mange av artene kan ligne på små maur.
Utseende Knøttsmå til små biller (gjerne ca. 1,5 mm), som oftest forholdsvis slanke og blanke, nakne eller med oppstående hår. Denne gruppen kjennes vanligvis lett fra andre perlebiller på at pronotum er forholdsvis stort og bredt, vanligvis tydelig bredere enn dekkvingene. Hodet er lite og mer eller mindre rundt, og skilt fra brystet med en tydelig "hals". Fasettøynene er små og utstående med ganske få fasetter, noen arter mangler fasettøyne. Antennene er ganske lange og kraftige, trådformede eller svakt kølleformede, og består av 11 ledd. Palpene er ofte forholdsvis lange og kølleformede. Brystskjoldet (pronotum) er vanligvis omtrent så langt som bredt og mye smalere enn dekkvingene. Ofte har det en tverr-rekke av dype groper nær roten. Dekkvingene dekker vanligvis hele bakkroppen. De er ofte ovale på form og bredest omkring midten. Hos mange arter har de noen dype groper ved roten. Beina er forholdsvis lange med fortykkede lår, ellers slanke. Fremlåret er ofte kledt med "sugekopper" som hjelper dem holde fast på byttet. Føttene er fem-leddete.
Levevis Perlebillene er rovdyr som stort sett lever av midd, spesielt pansermidd (Oribatida). Disse har et meget hardt skall og perlebillene har organer på frambeina for å holde byttet fast, og munndeler som passer til å borre eller sage seg gjennom middenes skall. Billene er ganske vanlige på litt fuktige steder, som i løvstrø, under steiner og i råtten, død ved. En god del arter lever i maur- eller termitt-tuer.
Fossile arter På grunn av sin beskjedne kroppsstørrelse blir disse billene lett bevart i rav. Det er derfor kjent relativt mange fossile arter i denne gruppen, og gruppens evolusjonære historie er bedre dokumentert enn de fleste andre biller.
Systematisk inndeling
Ordenen biller, Coleoptera Underordenen Polyphaga Overfamilien åtsel- og rovbiller, Staphylinoidea Familien kortvinger, Staphylinidae Latreille, 1802 Underfamilien perlebiller, Scydmaeninae Leach, 1815 Gruppe Cephenniitae Reitter, 1882 Stammen Cephenniini Reitter, 1882 Slekten Cephennodes Müller & Kunze, 1822 - ca. 170 arter Liste over artene
Cephennodes abdominalis Jałoszyński, 2007Jałoszyński, P. (2007) The Cephenniini of China. III. Cephennodes Reitter of Sichuan and Shaanxi (Coleoptera: Scydmaenidae). Genus, 18 (2): 151-207. Cephennodes akane Hoshina, 2020 Cephennodes alius Jałoszyński, 2025Paweł Jałoszyński (2025) The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101. DOI: 10.11646/zootaxa.5664.1.1 Cephennodes aliusmodi Jałoszyński, 2025 Cephennodes annapurnaensis Jałoszyński, 2007 Cephennodes antennatus Jałoszyński & Nomura, 2009Jałoszyński, P. & S. Nomura (2009) The Cephenniini (Coleoptera, Scydmaenidae) of Vietnam. Bulletin of the National Museum of Nature and Science (A: Zoology) 35 (3): 167-225. Cephennodes apicalis Jałoszyński & Nomura, 2009 Cephennodes araiorum (Jałoszyński, 2003)Jałoszyński, P. (2003) Chelonoidum araiorum sp. nov., an unusual species of the Cepheniini (Coleoptera, Scydmaenidae) from the intertidal zone of the Pacific coast of central Honshu, Japan. Bulletin of the National Science Museum (A: Zoology) 29 (4): 225-228. Cephennodes arcuatus Jałoszyński, 2025 Cephennodes aschnae Makhan, 2005Makhan, D. (2005) Four new Scydmaenidae species from Mount Jinyun, China (Coleoptera). Calodema 3: 6-13. Cephennodes ascipenis Jałoszyński, 2007 Cephennodes astoni Jałoszyński, 2011Jałoszyński, P. (2011) New records and new species of Cephennodes Reitter from Hong Kong (Coleoptera: Staphylinidae: Scydmaeninae). Genus 22 (1): 1-8. Cephennodes asymmetricus Jałoszyński, 2025 Cephennodes atewaensis Jałoszyński, 2010Jałoszyński, P. (2010) Afrotropical Cephenniini: Redescriptions of Cephennodes described by Schaufuss and Cauchois and three new species from Ghana, Nigeria and Uganda (Coleoptera: Staphylinidae: Scydmaeninae). Genus 21 (2): 175-190. Cephennodes atuin Jałoszyński & Nomura, 2009 Cephennodes baculifer Jałoszyński, 2007 Cephennodes bagmatianus Jałoszyński, 2017Jałoszyński, P. (2017) Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4349 (1): 1–120. Cephennodes bagongsilangensis Jałoszyński, 2009 Cephennodes bahrabisensis Jałoszyński, 2017 Cephennodes bakeri Blattný, 1929Blattný, C. (1929) Fauna Philippinica, Scydmaenidae, Coleoptera. Philippine Journal of Science 38 (3): 303-323., pls. 1-2. Cephennodes basilewskyi Besuchet, 1962Besuchet, C. (1962) Coleoptera Scydmaenidae: Cephenniini. Mission zoologique de l'I.R.S.A.C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXII. Annales du Musée Royal de l'Afrique Centrale, Tervuren (Série 8º: Sciences Zoologiques) 107: 420-423. Cephennodes bataanus Jałoszyński, 2009Jałoszyński, P. (2009) Cephenniini of the Philippines. Part 2. Two new species of Cephennodes Reitter from Mindanao (Coleoptera: Staphylinidae). Genus 20 (3): 391-397. Cephennodes besucheti (Kurbatov, 1995)Kurbatov, S. A. (1995) Sur les Euthiini et Cephenniini (Coleoptera, Scydmaenidae) de l'Extrême-Est de la Russie et du Japon. Revue Suisse de Zoologie 102 (4): 943-959. Cephennodes bicavatus Jałoszyński, 2015Jałoszyński, P. (2015) The Cephenniini of China. VI. New species and new records of Cephennodes Reitter from Hainan, Guangxi and Guangdong (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3990 (2): 221-234. Cephennodes bicolor Jałoszyński, 2007 Cephennodes bicribratus Jałoszyński, 2016Jałoszyński, P. (2016) The Cephenniini of China. VIII. New species and new records of Cephennodes Reitter of Hunan, Jiangxi, Zhejian and Fujian (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4079 (4): 415–428. Cephennodes biimpressus Jałoszyński, 2025 Cephennodes bilaminatus Jałoszyński, 2017 Cephennodes bilobatus Jałoszyński, 2017 Cephennodes bos Jałoszyński & Nomura, 2009 Cephennodes brachyclavatides Jałoszyński, 2026 Cephennodes brachyclavatus Jałoszyński, 2017 Cephennodes brachycornis Jałoszyński, 2025 Cephennodes brachylinguis Jałoszyński, 2007 Cephennodes brachyspinosus Jałoszyński, 2016 Cephennodes brinchangensis Jałoszyński, 2008Jałoszyński, P. (2008) A new bizarre species of Cephennodes from the Malay Peninsula (Coleoptera: Scydmaenidae). Genus 19 (1): 27-31. Cephennodes bubalus Jałoszyński & Nomura, 2009 Cephennodes buceros Jałoszyński & Nomura, 2009 Cephennodes bundibugyoensis Jałoszyński, 2010 Cephennodes capra Jałoszyński, 2012Jałoszyński, P. (2012) The Cephenniini of China. V. Cephennodes Reitter of Guizhou and Zhejiang (Coleoptera: Staphylinidae: Scydmaeninae). Genus 23 (2): 229-248. Cephennodes capricornis Jałoszyński, 2012 Cephennodes carinifrons Jałoszyński, 2007 Cephennodes caudatus Jałoszyński, 2007 Cephennodes cavatus Jałoszyński, 2025 Cephennodes cavifrons Jałoszyński, 2017 Cephennodes churtanus Jałoszyński, 2017 Cephennodes clavatus (Marsh, 1957Marsh, G. A. (1957) Family Scydmaenidae, pp. 271-280. I: Melville H. Hatch, The Beetles of the Pacific Northwest. Part II. Staphyliniformia. University of Washington Publications in Biology, 16 (2): ix + 384 pp. Cephennodes clavodentatus Jałoszyński, 2017 Cephennodes clypeatus Jałoszyński, 2012 Cephennodes clypeicornis Jałoszyński, 2012 Cephennodes concinnus (Marsh, 1957 Cephennodes cooteri Jałoszyński, 2007 Cephennodes cordilaminatus Jałoszyński, 2017 Cephennodes corporosus (LeConte, 1852) Cephennodes corpulentus Jałoszyński, 2015 Cephennodes cryptointricatus Jałoszyński, 2015 Cephennodes curvitibia Jałoszyński, 2025 Cephennodes datianpingensis Jałoszyński, 2016 Cephennodes dehongdaianus Jałoszyński, 2025 Cephennodes deinonychus Jałoszyński, 2025 Cephennodes depressifrons Jałoszyński & Nomura, 2009 Cephennodes deviator Jałoszyński, 2025 Cephennodes dilatatus Jałoszyński, 2025 Cephennodes dolakhanus Jałoszyński, 2017 Cephennodes elegantissimus Jałoszyński, 2015 Cephennodes elevatus Jałoszyński, 2015 Cephennodes elongatus Jałoszyński & Nomura, 2009 Cephennodes elytratus Jałoszyński, 2007 Cephennodes excavatus Jałoszyński, 2007 Cephennodes falcipenis Jałoszyński & Nomura, 2008 Cephennodes festivus (Schaufuss, 1889) Cephennodes frontocarinatus Jałoszyński & Nomura, 2008 Cephennodes frontorhinus Jałoszyński, 2025 Cephennodes furcatus Jałoszyński, 2017 Cephennodes gaozhaianus Jałoszyński, 2015 Cephennodes ghorepanianus Jałoszyński, 2017 Cephennodes glabella Castellini, 2007 Cephennodes glabrifrons Jałoszyński, 2025 Cephennodes gokarnaensis Jałoszyński, 2017 Cephennodes gopala Jałoszyński, 2017 Cephennodes gorkha Jałoszyński, 2017 Cephennodes gracilis Jałoszyński, 2025 Cephennodes graeseri Reitter, 1887 Cephennodes guadunanus Jałoszyński, 2007 Cephennodes guizhouanus Jałoszyński, 2012 Cephennodes gutta Jałoszyński, 2015 Cephennodes hamatus Jałoszyński, 2007 Cephennodes hellenicus (Roubal, 1913) Cephennodes hippopotamus Jałoszyński, 2007 Cephennodes hongkongensis Jałoszyński, 2008 Cephennodes huapingensis Jałoszyński, 2015 Cephennodes hystrix Jałoszyński, 2016 Cephennodes impressibos Jałoszyński, 2016 Cephennodes impressifrons Jałoszyński, 2007 Cephennodes indifferens Besuchet, 1962 Cephennodes inflaticornis Jałoszyński, 2017 Cephennodes inflatipes Jałoszyński, 2007 Cephennodes intricatus Jałoszyński, 2015 Cephennodes japonicus (Sharp, 1886) Cephennodes jizushanus Jałoszyński, 2025 Cephennodes kalopanianus Jałoszyński, 2017 Cephennodes karnaliensis Jałoszyński, 2017 Cephennodes koko Hoshina, 2021 Cephennodes kopeipes Jałoszyński, 2007 Cephennodes kotori Hoshina, 2021 Cephennodes kusunda Jałoszyński, 2017 Cephennodes lalitpuranus Jałoszyński, 2017 Cephennodes langbianganus Jałoszyński & Nomura, 2009 Cephennodes lateralis Jałoszyński & Nomura, 2009 Cephennodes latus (Motschulsky, 1851) Cephennodes leai Jałoszyński, 2008 Cephennodes leleupi Besuchet, 1962 Cephennodes leonensis Castellini, 2010 Cephennodes letheanus Jałoszyński, 2017 Cephennodes licchavi Jałoszyński, 2017 Cephennodes linguatoides Jałoszyński, 2025 Cephennodes linguatus Jałoszyński, 2025 Cephennodes loebli (Kurbatov, 1995) Cephennodes longicornis Jałoszyński, 2025 Cephennodes longilinguis Jałoszyński, 2007 Cephennodes longipes Jałoszyński, 2007 Cephennodes longulus Jałoszyński & Nomura, 2009 Cephennodes lucipetus Jałoszyński, 2013 Cephennodes lustricollis Jałoszyński & Nomura, 2009 Cephennodes lustrifrons Jałoszyński, 2007 Cephennodes luzonicus Blattný, 1929 Cephennodes magnus Jałoszyński, 2007 Cephennodes mahisapala Jałoszyński, 2017 Cephennodes malla Jałoszyński, 2017 Cephennodes malleiphallus Jałoszyński, 2007 Cephennodes manangensis Jałoszyński, 2017 Cephennodes mangmayanus Jałoszyński, 2017 Cephennodes marginatus Besuchet, 1962 Cephennodes martensi Jałoszyński, 2017 Cephennodes masuanus Jałoszyński, 2007 Cephennodes maya Hoshina, 2021 Cephennodes medusa Jałoszyński, 2015 Cephennodes meredaranus Jałoszyński, 2017 Cephennodes microphthalmus Jałoszyński, 2007 Cephennodes microsoma Jałoszyński, 2025 Cephennodes micrus Jałoszyński, 2025 Cephennodes minisulcatus Jałoszyński, 2017 Cephennodes minusculus Jałoszyński & Nomura, 2009 Cephennodes moderatus (Kurbatov, 1995) Cephennodes molestus Jałoszyński, 2025 Cephennodes monofoveatus Jałoszyński, 2015 Cephennodes monolaminatus Jałoszyński, 2017 Cephennodes mtubatubanus Jałoszyński, 2013 Cephennodes multisaetosus Jałoszyński & Nomura, 2008 Cephennodes mussauinsularis Jałoszyński, 2017 Cephennodes mustacifrons Jałoszyński & Nomura, 2009 Cephennodes mustangensis Jałoszyński, 2017 Cephennodes myagdiensis Jałoszyński, 2017 Cephennodes nanaka Hoshina, 2019 Cephennodes nanos Jałoszyński & Nomura, 2009