Epitonium (nomeados, em inglês, wentletraps -pl.; nome derivado do alemão wendeltreppe, cujo significado, traduzido para o português, é "escada em espiral", devido à forma de suas conchas, e assim também denominadas staircase shells ou ladder shells -pl.; em português - PRT -, escalárias -pl.; em castelhano, escalas ou escalarias -pl.) é um gênero de moluscos gastrópodes, marinhos e carnívoros de cnidários, pertencente à família Epitoniidae. Foi classificado por Peter Friedrich Röding, em 1798; com sua espécie-tipo, Epitonium scalare, descrita no ano de 1758 por Lineu.
Descrição da concha Conchas em forma de torre, com espiral geralmente alta e coloração de branca a castanha em graus variados. Com abertura de circular a oval, sem canal sifonal e com o relevo de sua concha geralmente bem esculpido de lamelas ou costelas de crescimento, espaçadas, não sendo esta uma característica única deste gênero. Opérculo córneo e circular, com finas e poucas voltas espirais e com núcleo quase central.
Espécies de Epitonium
Epitonium abyssicola (Schepman, 1909) Epitonium acanthopleurum (Verco, 1906) Epitonium acapulcanum Dall, 1917 Epitonium aciculinum (Hinds, 1844) Epitonium actinariophilum (Masahito & Habe, 1976) Epitonium aculeatum (G. B. Sowerby II, 1844) Epitonium adjunctum (Jousseaume, 1912) Epitonium aequale (Thiele, 1925) Epitonium agitabile (Jousseaume, 1912) Epitonium alabiforme Kilburn, 1985 Epitonium alatum (G. B. Sowerby II, 1844) Epitonium albidum (d'Orbigny, 1842) Epitonium albolineatum (G. B. Sowerby II, 1844) Epitonium algerianum (Weinkauff, 1866) Epitonium alizonae (Melvill, 1912) Epitonium amathusium (Melvill & Standen, 1903) Epitonium amiculum Kilburn, 1985 Epitonium amicum (Jousseaume, 1894) Epitonium amplexum Nakayama, 2003 Epitonium anabathmos Kilburn, 1985 Epitonium angulatum (Say, 1831) Epitonium angulicinctum (de Boury, 1913) Epitonium angustum (Dunker, 1861) Epitonium antisoa (Iredale, 1936) Epitonium apiculatum (Dall, 1889) Epitonium aranea Bonfitto, 2017 Epitonium arcanum DuShane, 1979 Epitonium artimi (Jousseaume, 1912) Epitonium atomus (E. A. Smith, 1890) Epitonium audouini (Jousseaume, 1894) Epitonium aureomaculatum (Masahito & Habe, 1973) Epitonium auritum (G. B. Sowerby II, 1844) Epitonium austrocaledonicum (Montrouzier, 1859) Epitonium avalites (Jousseaume, 1912) Epitonium babylonia (Dall, 1889) Epitonium barissum (Iredale, 1936) Epitonium beachportense (Cotton & Godfrey, 1938) Epitonium bellastriatum (Carpenter, 1864) Epitonium bellicosum Hedley, 1907 Epitonium bengalense (E. A. Smith, 1899) Epitonium berryi (Dall, 1907) Epitonium bevdeynzerae Garcia E., 2001 Epitonium blaisei (Barnard, 1963) Epitonium bonaespei (Barnard, 1963) Epitonium bouryi (Jousseaume, 1894) Epitonium boutetorum Garcia, 2016 Epitonium brachyspeira Kilburn, 1985 Epitonium brevissimum (Seguenza, 1876) Epitonium bucknilli Powell, 1924 Epitonium bulbulum (G. B. Sowerby II, 1844) Epitonium caamanoi Dall & Bartsch, 1910 Epitonium calideum (Melvill & Standen, 1903) Epitonium californicum Dall, 1917 Epitonium callipeplum Dall, 1919 Epitonium candeanum (d'Orbigny, 1842) Epitonium candidissimum (Monterosato, 1877) Epitonium canna Dall, 1919 Epitonium carchedon (Iredale, 1936) Epitonium castum (G. B. Sowerby II, 1873) Epitonium catalinense Dall, 1917 Epitonium catanuense (G. B. Sowerby II, 1844) Epitonium celesti (Aradas, 1854) Epitonium cerdantum (Melvill & Standen, 1903) Epitonium championi Clench & R. D. Turner, 1952 Epitonium chiarae Bozzetti, 2007 Epitonium chinglinae Y.-C. Lee & W.-L. Wu, 1998 Epitonium christiani Bozzetti, 2008 Epitonium christyi (Iredale, 1936) Epitonium clathratulum (Kanmacher, 1798) Epitonium clathrus (Linnaeus, 1758) Epitonium clementinum (Grateloup, 1840) Epitonium climacotum Kilburn, 1985 Epitonium columba Kilburn, 1985 Epitonium columnella Dall, 1917 Epitonium commodum (E. A. Smith, 1890) Epitonium conjunctum (Yokoyama, 1922) Epitonium connexum (G. B. Sowerby II, 1844) Epitonium continens (Melvill & Standen, 1903) Epitonium cookeanum Dall, 1917 Epitonium cophinodes (Melvill, 1904) Epitonium coretum (Iredale, 1936) Epitonium corniculum Y.-C. Lee & C.-W. Huang, 2016 Epitonium coutieri (Jousseaume, 1912) Epitonium couturieri (de Boury, 1912) Epitonium crassicostatum Gittenberger & Gittenberger, 2005 Epitonium crassum (G. B. Sowerby II, 1844) Epitonium creberrimum (Hinds, 1844) Epitonium crebricostatum (Carpenter, 1864) Epitonium crispatum (Pease, 1868) Epitonium crypticocorona Kilburn, 1985 Epitonium cultellicostum (de Boury, 1913) Epitonium cumingii (Carpenter, 1856) Epitonium curvilineatum (G. B. Sowerby II, 1844) Epitonium dallianum (Verrill & S. Smith [in Verrill], 1880) Epitonium deflersi (Jousseaume, 1912) Epitonium deificum (Melvill & Standen, 1903) Epitonium denticulatum (G. B. Sowerby II, 1844) Epitonium dentiscalpium (R. B. Watson, 1883) Epitonium deschampsi Garcia, 2003 Epitonium dubium (G. B. Sowerby II, 1844) Epitonium duocamurum Y.-C. Lee, 2001 Epitonium durhamianum Hertlein & A. M. Strong, 1951 Epitonium eboreum (E. A. Smith, 1906) Epitonium eclecticum (Melvill & Standen, 1903) Epitonium elenense (G. B. Sowerby II, 1844) Epitonium elisae Bozzetti, 2007 Epitonium eltanini (Dell, 1990) Epitonium emiliae (Melvill & Standen, 1903) Epitonium emydonesus Dall, 1917 Epitonium erranum (Melvill & Standen, 1896) Epitonium erroneum (Tapparone Canefri, 1876) Epitonium eulita (Dall & Simpson, 1901) Epitonium eusculptum (G. B. Sowerby III, 1903) Epitonium eutaenium Dall, 1917 Epitonium evanidstriatum Zelaya & Güller, 2017 Epitonium eximiellum (Masahito, Kuroda & Habe, 1971) Epitonium eximium (A. Adams & Reeve, 1850) Epitonium extenuicostum (de Boury, 1913) Epitonium fabrizioi Pastorino & Penchaszadeh, 1998 Epitonium falconi Kilburn, 1985 Epitonium fasciatum (G. B. Sowerby II, 1844) Epitonium fauroti (Jousseaume, 1912) Epitonium ferminense L. G. Brown, 2019 Epitonium ferrugineum (Mörch, 1852) Epitonium ferussacii (Audouin, 1826) Epitonium finitimum (Monterosato, 1890) Epitonium fischeri (R. B. Watson, 1897) Epitonium foliaceicosta (d'Orbigny, 1842) Epitonium fractum Dall, 1927 Epitonium fragile (Hanley, 1840) Epitonium friabile (G. B. Sowerby II, 1844) Epitonium frielei (Dall, 1889) Epitonium fucatum (Pease, 1861) Epitonium fulvovittatum (Dautzenberg, 1890) Epitonium georgettinum (Kiener, 1838) Epitonium glabratum (Hinds, 1844) Epitonium gloriolum (Melvill & Standen, 1901) Epitonium godfreyi (Cotton, 1938) Epitonium goldsmithi (DuShane, 1988) Epitonium goniophorum (Melvill & Standen, 1903) Epitonium gracile (G. B. Sowerby II, 1844) Epitonium graciliconfusum Nakayama, 2000 Epitonium gradaticostatum (Fenaux, 1938) Epitonium gradatum (G. B. Sowerby II, 1844) Epitonium gradilis (Jousseaume, 1912) Epitonium graviarmatum A. Gittenberger & E. Gittenberger, 2005 Epitonium gravieri (Jousseaume, 1912) Epitonium grayi (Nyst, 1871) Epitonium habeli Dall, 1917 Epitonium hamatulae Preston, 1915 Epitonium hancocki DuShane, 1970 Epitonium harimaense Makiyama, 1924 Epitonium harpa (Jousseaume, 1912) Epitonium harpago Kilburn, 1985 Epitonium heloris (Iredale, 1936) Epitonium hemmesi DuShane, 1988 Epitonium hexagonum (G. B. Sowerby II, 1844) Epitonium hindsii (Carpenter, 1856) Epitonium hispidulum (Monterosato, 1874) Epitonium histricosum (Jousseaume, 1912) Epitonium hueti Bozzetti, 2011 Epitonium huffmani DuShane & McLean, 1968 Epitonium humerosum (G. B. Sowerby III, 1901) Epitonium humphreysii (Kiener, 1838) Epitonium idalium (Melvill, 1912) Epitonium ilariae Bozzetti, 2007 Epitonium illovoense (Barnard, 1963) Epitonium imperiale (G. B. Sowerby II, 1844) Epitonium inaequale (Thiele, 1925) Epitonium incurvatum (Fenaux, 1938) Epitonium indianorum (Carpenter, 1865) Epitonium indistinctum (G. B. Sowerby II, 1844) Epitonium inexpertum L. G. Brown & Weil, 1999 Epitonium innesi (Jousseaume, 1912) Epitonium instrictum (E. von Martens, 1904) Epitonium interstriatum (G. B. Sowerby III, 1905) Epitonium irregulare (G. B. Sowerby II, 1844) Epitonium ishimotoi (Masahito & Habe, 1976) Epitonium janae Cossignani, 2020 Epitonium jani Segers, Swinnen & De Prins, 2009 Epitonium japonicum (Dunker, 1861) Epitonium jickelii (Clessin, 1897) Epitonium jimpyae Kilburn, 1985 Epitonium jolyi (Monterosato, 1878) Epitonium jomardi (Audouin, 1826) Epitonium jousseaumei (de Boury, 1886) Epitonium juanitae Garcia, 2003 Epitonium jukesianum (Forbes, 1852) Epitonium kastoroae Garcia, 2003 Epitonium kazusense (Yokoyama, 1922) Epitonium kilburni Drivas & Jay, 1989 Epitonium kiyohimae Nakayama, 2000 Epitonium klunzingeri (Clessin, 1897) Epitonium koshimagani (Nakayama, 1991) Epitonium koyamai (Nakayama, 1995) Epitonium kraussi (Nyst, 1871) Epitonium krebsii (Mörch, 1875) Epitonium labeo (Jousseaume, 1912) Epitonium lachrymulum (Jousseaume, 1912) Epitonium lacrima Kilburn, 1985 Epitonium laidlawi (Melvill & Standen, 1903) Epitonium latifasciatum (G. B. Sowerby II, 1874) Epitonium latum Bozzetti, 2009 Epitonium leali Garcia, 2011 Epitonium liliputanum (A. Adams, 1861) Epitonium linctum (de Boury & Monterosato, 1890) Epitonium lineolatum (G. B. Sowerby II, 1844) Epitonium lowei (Dall, 1906) Epitonium luceo (DuShane, 1988) Epitonium lyra (G. B. Sowerby II, 1844) Epitonium macleani DuShane, 1970 Epitonium macromphalus E. A. Smith, 1910 Epitonium maestratii Garcia, 2003 Epitonium malayanum (Thiele, 1925) Epitonium malcolmense (Melvill, 1898) Epitonium malhaensis (Jousseaume, 1894) Epitonium maraisi Kilburn, 1985 Epitonium margarita (Jousseaume, 1912) Epitonium marmoratum (G. B. Sowerby II, 1844) Epitonium matthewsae Clench & R. D. Turner, 1952 Epitonium melior (Melvill & Standen, 1903) Epitonium mellissi (E. A. Smith, 1890) Epitonium melvilli (Schepman, 1909) Epitonium millecostatum (Pease, 1861) Epitonium mindoroense (G. B. Sowerby II, 1844) Epitonium minorum (Iredale, 1936) Epitonium minutia (Jousseaume, 1912) Epitonium minuticostatum (de Boury, 1912) Epitonium mirabile Bozzetti, 2008 Epitonium miserum (de Boury, 1913) Epitonium mitraeforme (G. B. Sowerby II, 1844) Epitonium moolenbeeki van Aartsen, 1996 Epitonium morassii Bonfitto, 2017 Epitonium mucronatum (Fenaux, 1943) Epitonium multicostatum (G. B. Sowerby II, 1844) Epitonium multistriatum (Say, 1826) Epitonium muricatum (Risso, 1826) Epitonium mzambanum Kilburn, 1985 Epitonium nanum (Jeffreys, 1884) Epitonium nautlae (Mörch, 1875) Epitonium nearense L. G. Brown, 2019 Epitonium nitidella (Dall, 1889) Epitonium nodai Nakayama, 2000 Epitonium novangliae (Couthouy, 1838) Epitonium obesum (G. B. Sowerby II, 1847) Epitonium obliquum (G. B. Sowerby II, 1844) Epitonium obtusum (G. B. Sowerby, 1844) Epitonium occidentale (Nyst, 1871) Epitonium octogonum (G. B. Sowerby III, 1905) Epitonium okezoko (Habe, 1961) Epitonium opeas Kilburn, 1985 Epitonium oppositum (de Boury, 1921) Epitonium optabile (A. Adams, 1861) Epitonium ossium Nakayama, 2000 Epitonium ovale (G. B. Sowerby II, 1844) Epitonium padangense (Thiele, 1925) Epitonium pallasi (Kiener, 1838) Epitonium pallidizonatum (Masahito, Kuroda & Habe, 1971) Epitonium parspeciosum (Iredale, 1929) Epitonium parvonatrix Kilburn, 1985 Epitonium pasiphaes (Melvill, 1912) Epitonium paumotense (Pease, 1868) Epitonium perangustum (de Boury, 1913) Epitonium perfoliatum (Thiele, 1925) Epitonium perimense (Jousseaume, 1912) Epitonium perlucidum (Jousseaume, 1912) Epitonium perlucidum (Fenaux, 1943) Epitonium philippinarum (G. B. Sowerby II, 1844) Epitonium philtatum (R. B. Watson, 1886) Epitonium phymanthi R. Robertson, 1994 Epitonium pigrum Garcia, 2011 Epitonium platypleurum (Verco, 1906) Epitonium polacia (Dall, 1889) Epitonium politum (G. B. Sowerby II, 1844) Epitonium porrectum (Hinds, 1843) Epitonium profundum Nakayama, 2000